Publications

Organic Spinach Seed Production Manual

This guide includes basic instructions needed for on-farm organic spinach seed production. Topics include basic spinach seed biology, climatic requirements, soil fertility requirements, planting and cultivation, flowering and pollination, isolation requirements, genetic maintenance and improvement, harvesting, and common diseases.

• Navazio, J. and M. Colley. 2007. Principals and Practices of Organic Spinach Seed Production in the Pacific Northwest. Organic Seed Alliance. Available at https://projects.sare.org/wp-content/uploads/1626spinach_seed_manual.pdf

Articles

• Formiga, A., V. Joshi, M. Colley. 2023. Challenges in the market for organic spinach seed. eOrganic article. Available at https://eorganic.org/node/35758

• Joshi, V., Shi, A.K. Mishra, H. Gill, J. DiPiazza. 2022. Genetic dissection of nitrogen induced changes in the shoot and root biomass of spinach. Scientific Reports. 2022 Aug 12;12(1):13751. doi: 10.1038/s41598-022-18134-7. PMID: 35962022; PMCID: PMC9374745. Available at https://www.nature.com/articles/s41598-022-18134-7

Conference Presentations

• Benedict Analin, Dalton Thompson, Ainong Shi, and Vijay Joshi, 2026. Genome-Wide Insights into Leaf Architecture and Pigment Composition in Organically Grown Spinach. Oral presentation at the 2026 Southern Region American Society for Horticultural Science Annual Meeting in Louisville, Kentucky, on 02/1/2026. https://srashs2026.sched.com/event/2DK0w/genome-wide-insights-into-leaf-...
• Benedict Analin, Dalton Thompson, Ainong Shi, and Vijay Joshi, 2025. Natural Variation and Genome-Wide Association Mapping of Photosynthesis Traits in USDA Spinach Germplasm Under Organic Production.  In Hortscience, Vol. 60, No. 7. 113 S West St, Ste 200, Alexandria, Va 22314-2851 USA: American Society for Horticultural Science. https://ashs2025.sched.com/event/247cC/orgh-1-natural-variation-and-geno...
• Benedict Analin and Vijay Joshi, 2025. Differential Impacts of Nitrogen on Circadian Transcriptional Rhythms in Spinach. In Hortscience (Vol. 60, No. 7). 113 S West St, Ste 200, Alexandria, Va 22314-2851 USA: American Society for Horticultural Science. https://ashs2025.sched.com/event/247aS/pnm-4-differential-impacts-of-nit...
• Ainong Shi, Haizheng Xiong, Vijay Joshi, Benedict Analin, Carlos A Avila, and Beiquan Mou. 2025. Genome-Wide Analysis for Nutritional Components in Spinach. Poster presentation at The Plant & Animal Genome Conference (PAG-32), P362. https://plan.core-apps.com/pag32/abstract/11d52799fbee2e951df7857ea48f2e...
• Vijay Joshi, Benedict Analin, Micaela Colley, Ainong Shi, and Alice Formiga, 2025. Genetic Studies on The Productivity Traits of Organic Spinach. Scientific Conference on EUCARPIA Breeding to meet environmental and societal challenges, Page 152. EUCARPIA 2025_Abstracts_e-Book_Final_LS.pdf. 
• Vijay Joshi, Ainong Shi, Benedict Analin, Micaela Colley, and Alice Formiga, 2024. Phenotypic landscape of the photosynthetic performance and seed productivity of spinach germplasm under organic condition. Poster presentation at the American Society for Horticultural Science (ASHS) Annual Conference in Hawaii, USA. https://cdn.ymaws.com/ashs.org/resource/resmgr/2024_conference/Abstracts...
• Sandeep Kaur Sran, Dalton Thompson, and Vijay Joshi, 2025. The biofortification of spinach (Spinacia oleracea) with zinc and its interaction with nitrogen dynamics. Oral presentation at the American Society of Horticultural Sciences (ASHS) Annual Conference, in New Orleans, Louisiana, USA. https://ashs2025.sched.com/event/247jt. 
• Sandeep Kaur Sran, Dalton Thompson, and Vijay Joshi, 2025. Leveraging the potential of Methylobacterium symbioticum to boost spinach productivity. Poster presentation. American Society of Horticultural Sciences (ASHS) Annual Conference in New Orleans, Louisiana, USA. https://ashs2025.sched.com/event/247wC.